Quillwort References

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Bock, W. 1962. A study of fossil Isoëtes. Journal of Paleontology 36: 53–59.

Bolin, J. F., C. L. Hartwig, P. W. Schafran, and S. Komarnytsky. 2018. Application of DNA flow cytometry to aid species delimitation in Isoëtes. Castanea 83(1): 38–47.

Boom, B. M. 1980. Intersectional hybrids in Isoëtes. American Fern Journal 70: 1–4.

Boom, B. M. 1982. Synopsis of Isoëtes in the southeastern United States. Castanea 47: 38–59.

Bray, R. D., P. W. Schafran, and L. J. Musselman. 2018. Interesting, provocative, and enigmatic: morphological observations on southeastern quillworts (Isoëtes, Isoëtaceae, Lycophyta). Castanea 83(2): 263–269.

Brunton, D. F. 2015. Key to the quillworts (Isoëtes: Isoëtaceae) of the southeastern United States. American Fern Journal 105: 86–100.

Brunton D. F. 2016. Flat rock quillwort, Isoëtes graniticola, a new lycophyte from the southeastern United States. Rhodora 118: 261–275.

Brunton D. F. 2019. Distribution and taxonomy of Isoëtes tuckermanii subsp. acadiensis, comb. nov. (Isoëtaceae) in North America. The Canadian Field-Naturalist 132(4): 360–367.

Brunton, D. F., and D. M. Britton. 1997. Appalachian quillwort (Isoëtes appalachiana, sp. nov.; Isoëtaceae), a new pteridophyte for the eastern United States. Rhodora 99: 118–133.

Brunton, D. F., and D. M. Britton. 1998. Isoëtes microvela (Isoëtaceae), a new quillwort from the coastal plain of the southeastern United States. Rhodora 100: 261–275.

Brunton, D. F., D. M. Britton, and W. C. Taylor. 1994. Isoëtes hyemalis, sp. nov. (Isoëtaceae): a new quillwort from the southeastern United States. Castanea 59: 12–21.

Brunton, D. F., and J. McNeill. 2015. Status, distribution, and nomenclature of Northern Quillwort, Isoëtes septentrionalis (Isoëtaceae) in Canada. The Canadian Field-Naturalist 129(2): 174–180.

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Caplan, C. A., and C. R. Werth. 2000a. Isozymes of the Isoëtes riparia complex, I. Genetic variation and relatedness of diploid species. Systematic Botany 25(2): 235–259.

Caplan, C. A., and C. R. Werth. 2000b. Isozymes of the Isoëtes riparia complex, II. Ancestry and relationships of polyploids. Systematic Botany 25(2): 260–280.

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Dodge, R. 1896. The Ferns and Fern Allies of New England. W.N. Clute & Co., Binghamton, New York, USA.

Dodge, R. 1897. A new quillwort. Botanical Gazette 23: 32–29.

Duff, R. J., and E. E. Schilling. 2000. The chloroplast genome structure of the vascular plant Isoëtes is similar to that of the liverwort Marchantia. American Fern Journal 90: 51–59.

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Eaton, A. A. 1900. The genus Isoëtes in New England. Fernwort Papers 2: 1–16.

Engelmann, G. 1877. About the oaks of the United States. Transactions of the Academy of Science of St. Louis 3: 395.

Engelmann, G. 1882. The genus Isoëtes in North America. Transactions of the Academy of Science of St. Louis 4: 358–390.

Engelmann, G., and G. D. Butler. 1878. The species of Isoëtes of the Indian Territory. Botanical Gazette 3: 1–2.

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Foster, A. S. and E. M. Gifford, Jr. 1974. Comparative morphology of vascular plants. W.H. Freeman and Company, San Francisco, California, USA.

Freund, F. 2016. Characterizing quantitative variation in the glossopodia of three western North American Isoëtes species. American Fern Journal 106: 87–115.

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Heafner K. D., and R. D. Bray. 2005. Taxonomic reassessment of North American granite outcrop Isoëtes species with emphasis on vegetative morphology and I. piedmontana (Pfeiffer) Reed sensu lato. Castanea 70(3): 204–221.

Hickey, R. J. 1986. Isoëtes megaspore surface morphology: nomenclature, variation, and systematic importance. American Fern Journal 76: 1–16.

Hickey, R. J. 1990. Studies of neotropical Isoëtes L.: I. Euphyllum, a new subgenus. Annals of the Missouri Botanical Garden 77: 239–245.

Hickey, R. J., S. I. Guttman, and W. H. Eshbaugh. 1989. Evidence for post-translational modification of triose phosphate isomerase (TPI) in Isoëtes (Isoëtaceae). American Journal of Botany 76: 215–221.

Hickey, R. J., C. Macluf, and W. C. Taylor. 2003. A re-evaluation of Isoëtes savatieri Franchet in Argentina and Chile. American Fern Journal 93(3): 126–136.

Hickey, R. J., W. C. Taylor, and N. T. Luebke. 1989. The species concept in Pteridophyta with special reference to Isoëtes. American Fern Journal 79: 78–89.

Hoot, S. B., N. S. Napier, W. C. Taylor. 2004. Revealing unknown or extinct lineages within Isoëtes (Isoëtaceae) using DNA sequences from hybrids. American Journal of Botany 91(6): 899–904.

Hoot, S. B., and W. C. Taylor. 2001. The utility of nuclear ITS, a LEAFY homolog intron, and chloroplast atpB-rbcL spacer region data in phylogenetic analyses and species delimitation in Isoëtes. American Fern Journal 91(3): 166–177.

Hoot, S. B., W. C. Taylor, and N. S. Napier. 2006. Phylogeny and biogeography of Isoëtes (Isoëtaceae) based on nuclear and chloroplast DNA sequence data. Systematic Botany 31(3): 449–460.

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Karol, K. G., K. Arumuganathan, J. L. Boore, A. M. Duffy, K. D. E. Everett, J. D. Hall, S. K. Hansen, et al. 2010. Complete plastome sequences of Equisetum arvense and Isoëtes flaccida: implications for phylogeny and plastid genome evolution of early land plant lineages. BMC Evolutionary Biology 10: 321.

Karrfalt, E. E. 1977. Substrate penetration by the corm of Isoëtes. American Fern Journal 67: 1–4.

Kott, L. S., and D. M. Britton. 1983. Spore morphology and taxonomy of Isoëtes in northeastern North America. Canadian Journal of Botany 61: 3140–3163.

Kristen, U., and M. Biedermann. 1981. Ultrastructure, origin, and composition of the protein bodies in the ligule of Isoëtes lacustris L. Annals of Botany 48: 655–663.

Kristen, U., G. Liebezeit, and M. Biedermann. 1982. The ligule of Isoëtes lacustris: ultrastructure, mucilage composition, and a possible pathway of secretion. Annals of Botany 49: 569–584.

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Larsén, E., and C. Rydin. 2016. Disentangling the phylogeny of Isoëtes (Isoetales), using nuclear and plastid data. International Journal of Plant Science 177(2): 157–174.

Luebke, N. T., and J. M. Budke. 2003. Isoëtes tennesseensis (Isoëtaceae), an octoploid quillwort from Tennessee. American Fern Journal 93: 184–190.

Luebke, N. T., and W. C. Taylor. 1985. Detection of North American Isoëtes hybrids. American Journal of Botany 72: 924.

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Musselman, L. J. 2001. Georgia quillworts. Tipularia 2–19.

Musselman, L. J. 2002. Ornamentation of Isoëtes (Isoëtaceae, Lycophyta) microspores. Botanical Review 68(4): 474–487.

Musselman, L. J., and J. P. Roux. 2002. Isoëtes toximontana (Isoëtaceae), a new quillwort with green megaspores from the northern cape of South Africa. Novon 12: 504–507.

Musselman, L. J., R. D. Bray, and D. A. Knepper. 1996. Isoëtes ×bruntonii (Isoëtes engelmannii × Isoëtes hyemalis), a new hybrid quillwort from Virginia. American Fern Journal 86: 8–15. Musselman, L. J., R. D. Bray, P. W. Schafran, and W. C. Taylor. 2014. Misconceptions about Isoëtes. Paper presented at Association of Southeastern Biologists meeting in Spartanburg, South Carolina, USA.

Musselman, L. J., D. A. Knepper, R. D. Bray, C. A. Caplen, and C. Ballou. 1995. A new Isoëtes hybrid from Virginia. Castanea 60: 245–254.

Musselman, L. J., W. C. Taylor, and R. D. Bray. 2001. Isoëtes mattaponica (Isoëtaceae), a new diploid quillwort from freshwater tidal marshes of Virginia. Novon 11: 200–204.

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Osborn, T. G. B. 1922. Some observations on Isoëtes drummondii, A. Br. Annals of Botany 36: 41–54.

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Pereira, J. B., P. H. Labiak, T. Stützel, and C. Schulz. 2017. Origin and biogeography of the ancient genus Isoëtes with focus on the Neotropics. Botanical Journal of the Linnean Society 185: 253–271.

Pereira, J. B., P. H. Labiak, T. Stützel, and C. Schulz. 2019. Nuclear multi-locus phylogenetic inferences of polyploid Isoëtes species (Isoëtaceae) suggest several unknown diploid progenitors and a new polyploid species from South America. Botanical Journal of the Linnean Society 189: 6–22.

Pereira, J. B., M. Mittelbach, and P. H. Labiak. 2015. Studies on chromosome number and spore size in Brazilian Isoëtes. American Fern Journal 105(3): 226–237.

Pfeiffer, N. E. 1922. Monograph of the Isoëtaceae. Annals of the Missouri Botanical Garden 9(2): 79–233.

Pigg, K. B. 2001. Isoetalean lycopsid evolution: from the Devonian to the present. American Fern Journal 91(3): 99–114.

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Reed, C. F. 1965. Isoëtes in southeastern United States. Phytologia 12: 369–400.

Reed, C. F. 1945. Some nomenclatural changes in the genus Isoëtes. American Fern Journal 35: 77–86.

Rosenthal, M. A., S. R. Rosenthal, G. Johnson, W. C. Taylor, E. A. Zimmer. 2014. Isoëtes viridimontana: a previously unrecognized quillwort from Vermont, USA. American Fern Journal 104(1): 7–15.

Rydin, C., and N. Wikström. 2002. Phylogeny of Isoëtes (Lycopsida): resolving basal relationships using rbcL sequences. Taxon 51: 83–89.

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Schafran, P. W., E. A. Zimmer, W. C. Taylor, and L. J. Musselman. 2018. A whole chloroplast genome phylogeny of diploid species of Isoëtes (Isoëtaceae, Lycopodiophyta) in the southeastern United States. Castanea 83(2): 224–235.

Schuettpelz, E., and S. B. Hoot. 2006. Inferring the root of Isoëtes: exploring alternatives in the absence of an acceptable outgroup. Systematic Botany 31:258–270.

Sharma, B. D., and R. Singh. 1984. The ligule in Isoëtes. American Fern Journal 74: 22–28.

Singhurst, J. R., A. E. Rushing, C. K. Hanks, and W. C. Holmes. 2011. Isoëtes texana (Isoëtaceae): a new species from the Texas Coastal Bend. Phytoneuron 2011-22: 1–6.

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Takamiya, M., M. Watanabe, and K. Ono. 1999. Biosystematic studies on the genus Isoëtes (Isoëtaceae) in Japan. II. Meiotic behavior and reproductive mode of each cytotype. American Journal of Botany 83(10): 1309-1322.

Taylor, W. A. 1993. Megaspore wall ultrastructure in Isoëtes. American Journal of Botany 80: 165–171.

Taylor, W. C., and R. J. Hickey. 1992. Habitat, evolution, and speciation in Isoëtes. Annals of the Missouri Botanical Garden 79: 613–622.

Taylor, W. C., A. R. Lekschas, Q. F. Wang, X. Liu, N. S. Napier, and S. B. Hoot. 2004. Phylogenetic relationships of Isoëtes (Isoëtaceae) in China as revealed by nucleotide sequences of the nuclear ribosomal ITS region and the second intron of a LEAFY homolog. American Fern Journal 94(4): 196–205.

Taylor, W. C., N. T. Luebke, D. M. Britton, R. J. Hickey, and D. F. Brunton. 1993. Isoëtaceae. In FNA Editorial Committee [eds.], Flora of North America, North of Mexico, vol. 2, 64–75. Oxford University Press, New York, New York, USA.

Taylor, W. C., N. T. Luebke, and M. B. Smith. 1985. Speciation and hybridization in North American quillworts. Proceedings of the Royal Society of Edinburgh 86B: 259–263.

Taylor, W. C., R. H. Mohlenbrock, and J. A. Murphy. 1975. The spores and taxonomy of Isoëtes butleri and I. melanopoda. American Fern Journal 65(2): 33–38.

Troia, A., J. B. Pereira, C. Kim, and W. C. Taylor. 2016. The genus Isoëtes (Isoëtaceae): a provisional checklist of the accepted and unresolved taxa. Phytotaxa 277(2): 101–145.

Turner, N. A., W. C. Taylor, S. Masi, and M. E. Stupen. 2005. Confirming dioecy in Isoëtes butleri. American Fern Journal 95(2): 85–87.

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Yang, T., and X. Liu. 2015. Comparing photosynthetic characteristics of Isoëtes sinensis Palmer under submerged and terrestrial conditions. Scientific Reports 5: 17783.

Yang, T., and X. Liu. 2016. Comparative transcriptome analysis of Isoëtes sinensis under terrestrial and submerged conditions. Plant Molecular Biology Reporter 34(1): 136–145.

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